Three Tampa Bay Buccaneers players -- all on the defensive side of the ball -- will start training camp on the Physically Unable to Perform list, the Tampa Bay Times' Rick Stroud reported Tuesday morning.

Defensive end Da'Quan Bowers, defensive tackle Amobi Okoye and linebacker Adam Hayward will be out for Friday's opening day of camp, according to Stroud.

From Stroud's report:

Bowers suffered a torn Achilles tendon during an off-season workout in May. The Bucs still are hopeful to get him back sometime during the regular-season.

Okoye, who was signed to provide depth for oft-injured defensive tackles Gerald McCoy and Brian Price, underwent arthrscopic knee surgery in June and missed the mandatory minicamp.

Hayward, a sixth-year veteran who can play all three linebacker positions, struggled with a foot injury much of last season.

Players on the PUP list, as Stroud notes, can come off at any time during the preseason, but if anyone is still on the list once the season begins, they must sit out for six games per NFL rules.